 Following the events of Gol and Maia, Jak, Daxter, Keira, and Samos have been trying to unviel the mystery of the Precursors through the use of a machine. However, pressing the wrong button sends them into a dystopian city, where Jak is taken prisoner by Crimzon Guards, workers for Baron Praxis, and Daxter is left to find a way to get his friend out. During the two years, Jak has been exposed to Dark Eco that was being tested. Thankfully, Daxter rescues Jak, though he was surprised to see what happened to his friend. Jak now must exact revenge against Baron Praxis, as well as uncover a new evil about to erupt.